The bit which attaches to the top of your fork stanchions, and the bottom of your steerer tube.
WCA, having seen your bike in person at Swinley, I can say fair enough to you for trying it, but it’s not something I’d want to do myself. I think if your forks were setup so that they bottom out on occasion, I can see the tyre actually hitting the crown of your forks possibly. Certainly were you to hit a bump hard, the wheel could possibly deflect by a few mm, which itself would mean the tyre could hit the arch on the forks as it was FAR less than a finger gap from the tyre to the arch, a few Rizla’s more like 😉
Seen a few pics of peoples bikes running different combinations of wheel sizes front and back on MTBR forums as it seems to be the “in thing” to be doing in the States right now. Fair enough, whatever works for the individual is cool by me. But I can only say I’d not do it on a conventional fork designed for use with 26″ wheels as the potential for disaster is too great. Probably don’t need to point out the possible outcomes of what might happen if they tyre hit the crown under full compression! The only forks that are ok for it really are forks like my Maverick SC32’s, which are upside down so no arch, fitted with a travel reduction kit so that the fork bottoms out earlier and the tyre can’t hit the crown under full compression as pointed out by singlespeedstu.
FWIW, my mate has got a Chameleon setup as a 69er, with rigid forks up front. He loves it! Not to my taste but we’re all different so it’s all good. Personally I’m a 26″ wheel man through and through, and like my wheels to be the same size both ends.