I have to take one seat out to get my Alpine Five 29er in with both wheels off and it lying flat, probably would fit upright with just the front off.
i also have the Skoda bike rack in the back, bit limited as it only fits Q/R forks, but my 26″ bike and roadie fit right in upright, I still like to take a seat out though.
I really need to make adapters for it to take 15mm forks.
with a bit of jiggling i have fitted three bike in the back.
Two 29ers and a 26″ HT.