At the end of a descent the heating up thing could be a factor, but not over four hours of riding.
When you say stiff, is the spring rate increasing (like there’s more air in) or is it just rougher? It could be that they are low on lubrication oil and during the ride this is being spread thinly about the fork so it’s not being lubricated any more. Perhaps if you hang your bike up the little bit of oil collects at the seal again where it can lubricate again.
It’s not the norm, so start with a service.
I serviced the forks again this time I tightened the bottom bolts with the fork fully compressed, effectively creating a vacuum in the lowers when extended; but the force of the spring overcomes it. My forks are a lot better now.
This is like having a longer and slightly stronger negative spring. I did a similar thing on my Marzocchis by letting the air out of the damper above the oil – it did improve sensitivitiy but on that fork and bike it made it a bit too soft early in the stroke which is where that set-up needs it.
But I’d be worried about drawing muck and water into the fork running them with a vacuum in the lower leg.