I think its partly related to height
If you read Brants justification of the 456 having longer seat stays he mentions it.
Basically tall people end up sitting closer to the rear wheel than shorter people. This is only slightly offset by the longer top tube This means the taller folk, like me are often fighting to keep the front wheel down, even sat on the nore of the saddle. Longer seat stays help with this (as will saddle further forward)
This is one reason that my Fs will climb much steeper stuff than my old hard tail. Its a fuel ex. The FSR I demoed was a disaster. Putting the seat up sticks your ass over the rear wheel, even more wheelie tastic than my old hard tail.
The first Mountain bike I road was 28×28 bottom gear (26 inch gear)
The first MTB I owned was 28×30 bottom gear (24.3 inch)
I now have 22×34 on mt FS (17 inch)
Thats quite a big change!